diff --git a/Changes b/Changes
index d03bdabcfb..802a799022 100644
--- a/Changes
+++ b/Changes
@@ -6,6 +6,11 @@ Improvements
 
 - USDScene : Added loading of ArnoldProcedural prims as Cortex ExternalProcedural objects.
 
+Fixes
+-----
+
+- USDScene : Fixed writing of animated camera parameters.
+
 10.6.4.0 (relative to 10.6.3.1)
 ========
 
diff --git a/contrib/IECoreUSD/src/IECoreUSD/CameraAlgo.cpp b/contrib/IECoreUSD/src/IECoreUSD/CameraAlgo.cpp
index 677c4b25d8..034e28d881 100644
--- a/contrib/IECoreUSD/src/IECoreUSD/CameraAlgo.cpp
+++ b/contrib/IECoreUSD/src/IECoreUSD/CameraAlgo.cpp
@@ -156,27 +156,27 @@ bool writeCamera( const IECoreScene::Camera *camera, const pxr::UsdStagePtr &sta
 	auto usdCamera = pxr::UsdGeomCamera::Define( stage, path );
 	if( camera->getProjection() == "orthographic" )
 	{
-		usdCamera.GetProjectionAttr().Set( pxr::UsdGeomTokens->orthographic );
+		usdCamera.GetProjectionAttr().Set( pxr::UsdGeomTokens->orthographic, time );
 
 		// For ortho cameras, USD uses aperture units of tenths of scene units
-		usdCamera.GetHorizontalApertureAttr().Set( 10.0f * camera->getAperture()[0] );
-		usdCamera.GetVerticalApertureAttr().Set( 10.0f * camera->getAperture()[1] );
-		usdCamera.GetHorizontalApertureOffsetAttr().Set( 10.0f * camera->getApertureOffset()[0] );
-		usdCamera.GetVerticalApertureOffsetAttr().Set( 10.0f * camera->getApertureOffset()[1] );
+		usdCamera.GetHorizontalApertureAttr().Set( 10.0f * camera->getAperture()[0], time );
+		usdCamera.GetVerticalApertureAttr().Set( 10.0f * camera->getAperture()[1], time );
+		usdCamera.GetHorizontalApertureOffsetAttr().Set( 10.0f * camera->getApertureOffset()[0], time );
+		usdCamera.GetVerticalApertureOffsetAttr().Set( 10.0f * camera->getApertureOffset()[1], time );
 	}
 	else if( camera->getProjection() == "perspective" )
 	{
-		usdCamera.GetProjectionAttr().Set( pxr::UsdGeomTokens->perspective );
+		usdCamera.GetProjectionAttr().Set( pxr::UsdGeomTokens->perspective, time );
 
 		// We store focalLength and aperture in arbitary units.  USD uses tenths
 		// of scene units
 		float scale = 10.0f * camera->getFocalLengthWorldScale();
 
-		usdCamera.GetFocalLengthAttr().Set( camera->getFocalLength() * scale );
-		usdCamera.GetHorizontalApertureAttr().Set( camera->getAperture()[0] * scale );
-		usdCamera.GetVerticalApertureAttr().Set( camera->getAperture()[1] * scale );
-		usdCamera.GetHorizontalApertureOffsetAttr().Set( camera->getApertureOffset()[0] * scale );
-		usdCamera.GetVerticalApertureOffsetAttr().Set( camera->getApertureOffset()[1] * scale );
+		usdCamera.GetFocalLengthAttr().Set( camera->getFocalLength() * scale, time );
+		usdCamera.GetHorizontalApertureAttr().Set( camera->getAperture()[0] * scale, time );
+		usdCamera.GetVerticalApertureAttr().Set( camera->getAperture()[1] * scale, time );
+		usdCamera.GetHorizontalApertureOffsetAttr().Set( camera->getApertureOffset()[0] * scale, time );
+		usdCamera.GetVerticalApertureOffsetAttr().Set( camera->getApertureOffset()[1] * scale, time );
 	}
 	else
 	{
@@ -189,9 +189,9 @@ bool writeCamera( const IECoreScene::Camera *camera, const pxr::UsdStagePtr &sta
 		);
 	}
 
-	usdCamera.GetClippingRangeAttr().Set( pxr::GfVec2f( camera->getClippingPlanes().getValue() ) );
-	usdCamera.GetFStopAttr().Set( camera->getFStop() );
-	usdCamera.GetFocusDistanceAttr().Set( camera->getFocusDistance() );
+	usdCamera.GetClippingRangeAttr().Set( pxr::GfVec2f( camera->getClippingPlanes().getValue() ), time );
+	usdCamera.GetFStopAttr().Set( camera->getFStop(), time );
+	usdCamera.GetFocusDistanceAttr().Set( camera->getFocusDistance(), time );
 
 	if( camera->hasShutter() )
 	{
@@ -201,8 +201,8 @@ bool writeCamera( const IECoreScene::Camera *camera, const pxr::UsdStagePtr &sta
 		/// and Houdini plugin sources, I've been unable to find evidence for
 		/// anyone else doing this though, so maybe it's one of those things
 		/// everyone is just getting wrong?
-		usdCamera.GetShutterOpenAttr().Set( (double)camera->getShutter()[0] );
-		usdCamera.GetShutterCloseAttr().Set( (double)camera->getShutter()[1] );
+		usdCamera.GetShutterOpenAttr().Set( (double)camera->getShutter()[0], time );
+		usdCamera.GetShutterCloseAttr().Set( (double)camera->getShutter()[1], time );
 	}
 
 	return true;
diff --git a/contrib/IECoreUSD/test/IECoreUSD/USDSceneTest.py b/contrib/IECoreUSD/test/IECoreUSD/USDSceneTest.py
index af7902d4a1..369a6aed25 100644
--- a/contrib/IECoreUSD/test/IECoreUSD/USDSceneTest.py
+++ b/contrib/IECoreUSD/test/IECoreUSD/USDSceneTest.py
@@ -1306,7 +1306,7 @@ def formatCameraName( **kw ) :
 		for name, cortexCam in testCameras.items() :
 
 			cG = pxr.UsdGeom.Camera.Get( usdFile, "/" + name )
-			c = cG.GetCamera()
+			c = cG.GetCamera( 0.0 )
 
 			self.assertEqual( c.projection.name.lower(), cortexCam.getProjection() )
 
@@ -1329,8 +1329,8 @@ def formatCameraName( **kw ) :
 			self.assertEqual( c.fStop, cortexCam.getFStop() )
 			self.assertEqual( c.focusDistance, cortexCam.getFocusDistance() )
 			if cortexCam.hasShutter() :
-				self.assertEqual( cG.GetShutterOpenAttr().Get(), cortexCam.getShutter()[0] )
-				self.assertEqual( cG.GetShutterCloseAttr().Get(), cortexCam.getShutter()[1] )
+				self.assertEqual( cG.GetShutterOpenAttr().Get( 0.0 ), cortexCam.getShutter()[0] )
+				self.assertEqual( cG.GetShutterCloseAttr().Get( 0.0 ), cortexCam.getShutter()[1] )
 			else :
 				self.assertFalse( cG.GetShutterOpenAttr().HasAuthoredValue() )
 				self.assertFalse( cG.GetShutterCloseAttr().HasAuthoredValue() )
@@ -1426,6 +1426,34 @@ def assertVectorsAlmostEqual( a, b,**kw ) :
 
 			self.assertEqual( camera, roundTripCamera )
 
+	def testWriteAnimatedCamera( self ) :
+
+		fileName = os.path.join( self.temporaryDirectory(), "test.usda" )
+		root = IECoreScene.SceneInterface.create( fileName, IECore.IndexedIO.OpenMode.Write )
+		child = root.createChild( "camera" )
+
+		cameras = {}
+		for time in range( 0, 10 ) :
+
+			camera = IECoreScene.Camera()
+			camera.setProjection( "perspective" )
+			camera.setShutter( imath.V2f( time - 0.25, time + 0.25 ) )
+			camera.setFocalLength( 1.0 + time * 0.1 )
+			camera.setClippingPlanes( imath.V2f( time, 1000 ) )
+
+			child.writeObject( camera, time )
+			cameras[time] = camera
+
+		del child, root
+
+		root = IECoreScene.SceneInterface.create( fileName, IECore.IndexedIO.OpenMode.Read )
+		child = root.child( "camera" )
+
+		for time in range( 0, 10 ) :
+			camera = child.readObject( time )
+			for parameter in ( "shutter", "focalLength", "clippingPlanes" ) :
+				self.assertEqual( camera.parameters()[parameter], cameras[time].parameters()[parameter] )
+
 	def testCornersAndCreases( self ) :
 
 		mesh = IECoreScene.MeshPrimitive.createPlane( imath.Box2f( imath.V2f( -1 ), imath.V2f( 1 ) ) )